Ajankohtaista

Doctoral Researchers – University of Helsinki

5.5.2026 | Avoimet työpaikat

Several positions are open! Check further information from the links below.

1. Doctoral Researcher (MSCA), Generation and optimization of oncolytic viruses for treatment of GBM

    https://jobs.helsinki.fi/job/Helsinki-Doctoral-Researcher-(MSCA),-Generation-and-optimization-of-oncolytic-viruses-for-treatment-of-GBM/1359244157/?utm_source=duunitori.fi&utm_medium=referral

    2. Doctoral Researcher (MSCA), Generation and tracking of armed rADV with imaging reporter genes

    https://jobs.helsinki.fi/job/Helsinki-Doctoral-Researcher-(MSCA),-Generation-and-tracking-of-armed-rADV-with-imaging-reporter-genes/1359243957/?utm_source=duunitori.fi&utm_medium=referral

    3. Doctoral Researcher (MSCA), Assessing the Immunological Response of Oncolytic Virus Nanoplatforms

    https://jobs.helsinki.fi/job/Helsinki-Doctoral-Researcher-%28MSCA%29%2C-Assessing-the-Immunological-Response-of-Oncolytic-Virus-Nanoplatforms/1359244257/?feedId=350602&utm_source=CareerSite_UniversityOfHelsinki&utm_medium=referral

    Apply latest by 28.5.2026

    Doctoral Researcher, Cancer Data Science – University of Helsinki

    5.5.2026 | Avoimet työpaikat

    https://jobs.helsinki.fi/job/Helsinki-Doctoral-Researcher%2C-Cancer-Data-Science/1358307857/?feedId=350602&utm_source=CareerSite_UniversityOfHelsinki&utm_medium=referral

    Apply latest by 31.5.2026

    University of Helsinki, Faculty of Medicine invites applications for the position of 

    Doctoral Researcher 

    in the Cancer Data Science group led by Associate Professor Mariike Kuijjer, at the iCAN Flagship in Digital Precision Cancer Medicine at the Faculty of Medicine, University of Helsinki. 

    The position is funded through a project by the Sigrid Jusélius Foundation titled “Regulatory determinants of metastatic progression in high-grade serous ovarian cancer.” The ideal starting date is summer or early fall 2026, but this is flexible and can be discussed. The appointment is a full-time position and is made for a period of three years with possible extension depending on future funding.


    Our research

    The Kuijjer group was established in 2018 at the University of Oslo and expanded to the University of Helsinki in 2025. The group focuses on developing computational and data science approaches to increase our understanding of gene regulation and how rewiring of gene regulatory networks can influence cancer development, progression, and disease heterogeneity. The group’s current projects use network science and artificial intelligence to develop tools that (i) integrate multi-modal data, (ii) model gene regulation at increased genomic and cellular resolution, (iii) refine the analysis of genome-wide regulatory networks. The group applies these tools to study various cancer types, including ovarian cancer. For more information, visit kuijjerlab.org.


    About the role

    The advertised project aims to identify regulatory determinants of ovarian cancer progression through regulatory network modeling and analysis. The selected candidate will apply and extend network modeling approaches developed in the Kuijjer group to model patient- and cell type-specific gene regulatory networks using bulk and single-cell ovarian cancer data. These networks will be analyzed to define patient-specific regulatory heterogeneity in primary ovarian cancer, identify regulatory rewiring that drives metastatic progression, characterize the regulatory mechanisms modulated by therapeutic pressure, and assess how these programs relate to therapy response. The project will leverage various publicly available datasets as well as data from the in-house Oncosys-OVA study led by our collaborator Prof. Anniina Färkkilä, University of Helsinki. Overall, the project combines methodological development with biological discovery, offering opportunities to contribute both to computational innovation and to mechanistic insights into ovarian cancer progression.


    The candidate

    We seek a highly motivated candidate with strong computational skills in the field of high-throughput omics data analysis, comparative genomics, functional genomics, or related fields. The candidate is excited about applying computational tools to answer questions in cancer research. They are collaborative and creative, and have strong programming skills dedicated to the analysis of large-scale genomics data. Experience with the analysis of omics data is required for this position. Knowledge of cancer biology is a strong advantage. Experience with high-performance computing, genomic data integration, biological network analysis, and/or gene regulation is desirable, but not required.

    The position is open to applicants with a Master’s degree in computational biology, bioinformatics, biostatistics, cancer genomics, network science, or related fields. Candidates who are finishing their Master’s thesis are also welcome to apply. They should mention the expected date of their defense and steps that need to be completed to finish their thesis in the cover letter.


    Qualification requirements

    • Master’s degree in computational biology, bioinformatics, biostatistics, cancer genomics, network science, or a related field. Applicants with a background in biology or (bio)medicine are welcome to apply, provided they have documented expertise in computational biology
    • Proficiency in programming (such as Python, R, Bash)
    • Experience with analysis of high-throughput omics data is required
    • Knowledge of cancer biology is a strong advantage
    • Experience with genomic data integration, biological network analysis, and/or gene regulation is an advantage
    • Experience with high-performance computing is desirable
    • Strong analytical and problem-solving skills
    • Teamwork skills and the willingness to collaborate, share ideas, and contribute to a multidisciplinary research environment spanning Helsinki and Oslo
    • Strong communication skills in English, with both written and verbal proficiency


    The appointee should either already have the right to pursue a doctoral degree at the University of Helsinki by the start of the appointment or apply for the right and obtain it within the probationary period of six months of their appointment. If the candidate does not already have the right to pursue a doctoral degree at the University of Helsinki, it must be applied for separately: www.helsinki.fi/en/research/doctoral-education/the-application-process-in-a-nutshell. The requirements for pursuing a doctoral degree at the University of Helsinki can be found at https://www.helsinki.fi/en/research/doctoral-education/eligibility-and-educational-documents.

    The chosen applicant is expected to reside in Finland while employed by the University of Helsinki.


    What we offer

    • A salary of  2600 – 2800 €/month depending on the candidate’s qualifications
    • A professional, stimulating working environment
    • The University of Helsinki also offers comprehensive services to its employees, including occupational health care and health insurance, unemployment and pension fund, a generous holiday package, sports facilities, and opportunities for professional development: https://www.helsinki.fi/en/about-us/careers.


    How to apply

    Applications should be submitted through the University of Helsinki Recruitment System via the ”Apply now” button.

    Internal applicants (i.e., current employees of the University of Helsinki) submit their applications by using the Employee Login button.

    The application must include:

    1. A cover letter, stating your motivation, scientific background, documented experience with large-scale genomic analyses, and research interests
    2. A detailed CV with a list of publications and other research output, if applicable
    3. 2-3 references (name, institution, e-mail, telephone number, and relation to the candidate)

    These should be uploaded in .pdf format. Applicants may also upload their Master’s thesis or other representative research work, if available. Please submit your application through the web-based recruitment system linked in the announcement. Although the recruitment system includes a basic CV, we ask candidates to also include a separate, detailed CV in .pdf format with their application. Applications without a cover letter and/or detailed CV will not be considered further. 

    The closing date for applications is 31.5.2026.

    Inquiries about the position can be directed to Mariike Kuijjer: mariike.kuijjer (at) helsinki.fi.

    If you need technical support with the recruitment system or additional information about the application process, please contact HelsinkiUni Help, uni-help (at) helsinki.fi. 

    Doctoral Researcher in Cell Biology, Centre of Excellence in Immune – Endothelial Interfaces – University of Helsinki

    5.5.2026 | Avoimet työpaikat

    https://jobs.helsinki.fi/job/Helsinki-Doctoral-Researcher-in-Cell-Biology,-Centre-of-Excellence-in-Immune-Endothelial-Interfaces/1358288757/?utm_source=duunitori.fi&utm_medium=referral

    Apply latest by 13.5.2026

    The newly established Finnish Centre of Excellence in Immune – Endothelial Interfaces (IMMENs) invites applications for a fixed-term (2-years) position of a

    Doctoral Researcher 

    The successful candidate will work in the research group of Dr. Kari Vaahtomeri at the Faculty of Medicine, University of Helsinki.

    Scientific context

    IMMENs, composed of research groups at the University of Helsinki (director Dr. Taija Mäkinen and Dr. Kari Vaahtomeri) and Åbo Akademi in Turku (Dr. Cecilia Sahlgren and Dr. Guillaume Jacquemet), investigates the role of lymphatic vessel endothelium in controlling immune responses. Current evidence shows that much of immune regulation occurs locally at the site of inflammation. Recent studies have revealed important crosstalk between lymphatic endothelium and immune cells, resulting in phenotypic changes that control immune cell trafficking, fate and function. IMMENs will develop novel experimental approaches to uncover the mechanisms and therapeutic potential of lymphatic endothelial immunomodulation in homeostasis, inflammation and tumorigenesis using in vivo genetic mouse models, advanced live and intravital imaging, engineered microchip models, primary cell co-culture systems and novel microscopy and analysis methods. In this context, Vaahtomeri laboratory addresses fundamental cell biology mechanisms of cell migration/interactions.

    Who are we looking for

    We are looking for candidates with a strong background in life sciences (for example, cell biology, biochemistry or mouse models). The candidate should be highly motivated to investigate key open questions in the fields of:

    • Cell Biology
    • Vascular biology
    • Immunology

    At the pre-doctoral level, previous experience in cell biology, vascular biology, mouse models, immunology or high-end imaging approaches is considered an advantage. 

    Doctoral Researcher must obtain the right to pursue doctoral studies at the University of Helsinki within the trial period of six months. Candidates should be able to demonstrate abstract thinking as well as the potential to develop and pursue scientific projects in the fields relevant for this call. Furthermore, the candidate should have excellent communication skills in English and the ability to work in an international environment. 

    What we offer

    The initial appointment is for 2 years and includes a possibility for extension upon success in doctoral research. The position will be filled immediately upon identification of suitable candidates. The starting date is negotiable. Salary will be commensurate with qualifications and experience based on the University of Helsinki guidelines. The chosen applicant is expected to reside in Finland while employed by the University of Helsinki. 

    The work is supervised by Dr. Kari Vaahtomeri and takes place in the context of the active research community of IMMENs groups. The research group has prominent international connections and is located at the medical campuses in the University of Helsinki. The immediate research environment provides state-of-the-art facilities, cutting-edge expertise in vascular-, tumor-, immunological- and stem cell biology, and great synergy with renowned research groups.

    More information on the IMMENS and Vaahtomeri research group can be found here: 
    https://www.helsinki.fi/en/researchgroups/immune-endothelial-interfaces 
    https://www.helsinki.fi/en/researchgroups/cell-communication


    How to apply

    Please submit your application as a single PDF file through the University of Helsinki Recruitment System by clicking the “Apply now” button. 

    Applicants who are already employees of the University of Helsinki are requested to submit their application via the ”Employee Login” button.

    Please attach the following documents as a single PDF file:

    • A cover letter that details your previous knowledge and research interests and motivation for applying this position
    • CV, including a possible list of publications
    • Transcript of study records
    • Contact details of at least two referees

    For more information on the position and specific projects, please contact Dr. Kari Vaahtomeri (kari.vaahtomeri (at) helsinki.fi).

    If you need technical support with the recruitment system or additional information about the application process, please contact HelsinkiUni Help, uni-help (at) helsinki.fi. 

    The closing date for the applications is on Wednesday the 13th of May 2026.

    Recent relevant publications:

    Liaqat et al. “Spatially targeted chemokine exocytosis guides transmigration at lymphatic endothelial multicellular junctions”. The EMBO Journal. 2024 Aug;43(15):3141-3174. doi: 10.1038/s44318-024-00129-x. 

    Can Ucar et al. ”Self-organized and directed branching results in space-filling coverage in developing dermal lymphatic networks”. Nat Commun. 2023 Sep 21;14(1):5878. doi: 10.1038/s41467-023-41456-7. 

    Project Researcher / Doctoral Researcher – Turku PET Centre

    18.1.2026 | Avoimet työpaikat

    https://ats.talentadore.com/apply/projektitutkija-vaitoskirjatutkija/m6rMgp?lang=en&UTUID=21181

    Apply latest by 30.1.2026

    Turku PET Centre is looking for a Project Researcher or Doctoral Researcher in PET imaging for a fixed-term position lasting approximately 2.5 years, starting no later than 1 March 2026 (exact start date negotiable), ending 31 August, 2028.  The position may be extended after this date. 

    We are looking for an enthusiastic and skilled researcher for our project, which involves developing new methods for modeling and reconstructing total body (large axial field of view) PET data. The project is led by Docent Jarmo Teuho and funded by the Academy of Finland. 

    Job description

    Your responsibility is the following: to develop kinetic modelling and new reconstruction methods for total body (large axial field of view) PET/CT, analyse and report data and participate as a member of the research group and conduct scientific publishing, supervision and reporting actions as part of your PhD and as a part of a larger research group.

    Who we are looking for

    We are looking for a candidate with an Medical Physics or Biomedical Engineering (or equivalent) background. Preferably you already have some experience from imaging methods, especially PET imaging.

    The position requires excellent programming skills with Python and MATLAB. Preferably, you are able to provide some proven examples. Additional programming skills and knowledge of GPU programming are a plus, but are not essential requirements.

    The essential skills we require are good written and spoken English skills, and the ability to communicate clearly, capability for independent research work and scientific publishing, as well as group work skills.

    A Doctoral Researcher must have a study right in the Turku University Doctoral Programme, otherwise the title of a Project Researcher is used. To be eligible for the post of Project Researcher, candidates must have a Master’s degree by the closing date for applications.

    The qualification requirements for doctoral researchers are set in the University of Turku Rules of Procedure

    We value collegiality and promote equality, diversity, and inclusivity. We encourage all qualified applicants with different backgrounds to apply for our open positions.

    Apply between 11 December 2025 and 30 January 2026 16:00 (Europe/Helsinki)

    Benefits

    We offer you responsible and interesting tasks and the opportunity to develop your professional skills in a versatile operating environment. You get a professional and inspiring team to support your work. We take care of your well-being at work and the functionality of your everyday life, e.g. with the help of flexible working hours and our well-being services.

    • Possibilities to develop your expertise – We encourage our personnel to develop their professional expertise by offering versatile training and competence development possibilities which employees can take advantage of during their working hours. Also, the course offering of Open University is free of charge for our personnel.
    • Your well-being is important to us! – In addition to comprehensive occupational health care, our own well-being services monitor, support and develop the well-being of our personnel. You are entitled to use the CampusSport’s services at a personnel price.
    • We support your work-life balance – Flexible working hours and extensive remote work options help you balance work and leisure time. 
    • An inspiring and supportive work community – We encourage each other and value working together!

    Please read more about University of Turku as an employee from our Come work with us! pages.

    The European Commission has awarded the University of Turku the right to use the HR Excellence in Research quality label. The label is a token of the University’s commitment to continuous development of the position and working conditions of researchers according to the guidelines set forth in the European Charter for Researchers. 

    The university offers good support and orientation for international hires. Please learn more about the Finnish culture and relocation to Finland:

    The University of Turku is among the top of the world’s universities in most rankings and among the best universities in the world in certain fields. Our high-quality basic research, a multidisciplinary research profile and close cooperation in research infrastructures creates the basis for international top research. Read more about University of Turku in international university rankings.

    The University of Turku has several high-quality research infrastructures required for cutting-edge research. Learn more about the roadmap of research infrastructures and how the research infrastructure and expert teams offer tailored solutions for businesses and organizations.

    Salary and Trial Period

    The salary is determined in accordance with the University salary system for teaching and research personnel. The salary will be based on demand levels 2-4, where the task-specific salary component is 2 281,48-2 844,66 euros per month. In addition, a personal work performance component will be paid.

    A six (6) month trial period applies to the position.

    How to apply

    Please submit your application through our online recruitment system. The closing date for applications is 30.1.2026 at 16.00 (Europe/Helsinki).

    Please write your application and accompanying documents in English. Provide the following documents as an attachement to your application:

    – Letter of motivation

    – Curriculum vitate

    – A copy of your degree certificates

    – A list of publications

    – Letter of recommendation from two previous supervisors with contact information

    Please be careful when filling out the application and make sure you submit all the required attachments.

    Further Information

    For further information about the position, please contact: Docent Jarmo Teuho, jarmo.teuho@utu.fi. 

    With questions about the application process, please contact: HR Specialist Mirjami Jaakkola, mirjami.a.jaakkola@utu.fi. 

    Doctoral Researcher, Molecular Medicine (Celiac Disease & Immunometabolism) – University of Oulu

    18.1.2026 | Avoimet työpaikat

    https://oulunyliopisto.varbi.com/fi/what:job/jobID:890108

    Apply latest by 31.1.2026

    Faculty of Biochemistry and Molecular Medicine, ECM and Hypoxia 

    The University of Oulu is a multidisciplinary, international research university, with about 4000 employees who produce new knowledge based on high-standards research and provide research based education to build a more sustainable, smarter, and more humane world. The University of Oulu community has about 17,000 people in total. Our northern scientific community operates globally and creates conditions for the emergence of innovations.  

    Doctoral Researcher

    to join the Intestinal Signaling and Epigenetics research group led by Associate Professor Keijo Viiri in the Faculty of Biochemistry and Molecular Medicine.

    This position is part of the HEALTH-DIMENSIONS (Profi 8) research programme, a national profiling action funded by the Research Council of Finland. The programme focuses on ”Multidimensional Big Data and Life-Course Health,” with a specific emphasis on molecular-level mechanistic validation of diseases.

    About the Job

    Think of the human body as a complex computer system. Celiac disease is like a software glitch triggered by an external input (dietary gluten) that causes the hardware (the intestine) to malfunction. This research project acts as a system diagnostic—examining the “code” (functional epigenetics) and the “power supply” (metabolism) to identify a “patch” (targeted therapy) that restores the system to its normal state. Above all, this project champions discovery science—uncovering fundamental principles that transcend disease-specific boundaries and generate knowledge with transformative potential across diverse health challenges.

    The successful candidate will contribute to an interdisciplinary research environment that bridges clinical data with fundamental molecular biology. The project aims to understand the root causes of intestinal damage and develop personalized treatment strategies for autoimmune conditions.

    Key research themes include:

    • Precision Immunotherapy: Dissecting molecular pathways driving Celiac Disease (CeD) and its refractory forms to unlock new therapeutic strategies.
    • Advanced Disease Modelling: Using omics technologies and patient-derived as well as transgenic murine models to understand why some patients continue to experience symptoms despite standard treatments.
    • Metabolism and Cell Health: Exploring how cellular metabolism and epigenetic gene regulation interact to maintain a healthy intestinal lining and how inflammatory responses disrupt this balance.
    • Viral Factors: Examining how external triggers, such as viral infections, may predispose individuals to autoimmune responses.

    What We Offer

    • A supportive, multidisciplinary team where you can grow as an expert in molecular medicine and functional epigenetics.
    • Access to cutting-edge research infrastructure, including organoid models and advanced sequencing technologies.
    • Wellness benefit ePassi covering sport, culture, and well-being. Read more about other staff benefits.  
    • Development and career options of the big organization.
    • Work that matters and a workplace that promotes flexibility and work-life balance. Read more about working with us.
    • Our Buddy Programme and Spouse Network support you and your close ones in settling into Oulu.
    • We have an HR Excellence in Research -quality label, which is a recognition awarded by the European Commission for the development of researchers’ working conditions and careers. 
    • Finland is one of the most livable countries in the world, with a high quality of life, safety, an excellent education system, and a competitive economy. Read more about living in Oulu.

    Who Are You?

    To succeed in this position, we require:

    • A Master’s degree in a relevant field (e.g., Molecular Medicine, Biochemistry, Cell Biology, or Health Sciences). The Master’s degree must have been completed before the start of the doctoral researcher’s work.
    • See also the requirements of the University of Oulu Graduate School (UniOGS)
    • Fluent oral and written English skills.
    • A strong interest in autoimmunity, metabolism, and epigenetic gene regulation.

    We also appreciate:

    • Prior experience in cell culture, molecular biology techniques, imaging and animal models.
    • Familiarity with bioinformatics or the analysis of large biological datasets.
    • A proactive attitude and the ability to work in an international research environment.

    Salary

    The position is fixed-term for three years. Salary is based on levels 2–4 of the demand level chart for teaching and research staff at Finnish universities. In addition, a personal performance component will be paid, with a typical starting gross salary between €2,600 and €2,900 per month. A trial period of 6 months is applied for the position.

    How to Apply?

    Interested? If yes, please apply by 31.1.2026 (23:59 Finnish local time) through our recruitment system.

    The application must be written in English and include the following:

    1. Cover letter summarizing your research interests and suitability.
    2. CV following the guidelines of the Finnish National Board on Research Integrity.
    3. Certificates/Diplomas.
    4. Contact information for two referees.

    The eligible applicants fitting best in the profile expected for the position will be invited to an on-site or remote interview. All applicants will be notified during the selection process.

    We welcome applicants from all backgrounds, such as people of different ages, genders, and linguistic, cultural, or minority groups. 

    Contact Information

    If you have any further questions, please contact Associate Professor Keijo Viiri, keijo.viiri@oulu.fi 

    Postdoctoral researcher and doctoral researcher in plant science and algae research- University of Turku

    4.11.2025 | Avoimet työpaikat

    Apply latest by 27.11.

    https://ats.talentadore.com/apply/tutkijatohtori-ja-vaitoskirjatutkija-kasvitieteeseen-ja-levatutkimukseen/mEovXW?lang=en&UTUID=21078

    We are seeking a Postdoctoral researcher and Doctoral researcher (or Research assistant) to join the Photosynthetic Microbes group led by Professor Yagut Allahverdiyeva-Rinne. The positions are part of the AlgaCircle project funded by Business Finland and several Finnish companies. The work will be carried out at the Faculty of Technology, within the Department of Life Technologies, in Turku, Finland. Employment will start on 1 January 2026, or as agreed. The successful candidate will be offered a fixed one-year contract, with the possibility of extension. Doctoral Researcher will be enrolled to the University of Turku Graduate School  (UTUGS) and after that the person will be employed for at least 3 years.

    About the Photosynthetic Microbes Research Group

    We study the regulation of photosynthesis and apply photosynthetic principles to develop sustainable biotechnologies. Our research encompasses several areas, including the elucidation of photosynthetic electron transport pathways, engineering photosynthesis for enhanced bioproduction of solar chemicals, biophotovoltaic (BPV) systems, and large-scale algal cultivation for phytoremediation and the production of green chemicals such as biostimulants and biopesticides

    The truly international and large PhotoMicrobes group is involved in several EU, Nordic, and national projects and regularly publishes its research outcomes in highly respected journals. We have established a broad research network with connections to leading international laboratories and industrial partners. Our team members are not only active participants in international conferences but also regularly organize major symposia and meetings in Turku (e.g., the Nordic Photosynthesis Congress, the International Conference on Renewable Resources 2025, and the Nordic Algal Biotechnology Conference).

    We are developing the national FIN-BioFoundry research infrastructure roadmap, which provides state-of-the-art equipment and methodologies. In addition, we are actively involved in science policymaking through the SYNERGY pan-European initiative.

    To learn more about the Photosynthetic Microbes research group, you can visit our website at www.utu.fi/photo-microbes or follow us on LinkedIN and BlueSky at @PhotoMicrobes.

    Job description

    The AlgaCircle project aims to utilize effluents from Finnish industries to cultivate microalgae and upgrade the resulting biomass through a multiproduct biorefinery approach. The project is carried out in collaboration with several industrial partners.

    Position #1 will focus on testing different algal formulations on crops and will work in close collaboration with researchers operating large photobioreactors. Part of the experiments will be conducted together with the NAPPI plant phenotyping platform.

    Position #2 will focus on optimizing the phytoremediation of industrial effluents by adjusting photobioreactor parameters and applying genetic engineering and synthetic biology approaches.

    Apply between 16 October 2025 and 27 November 2025 16:00 (Europe/Helsinki)

    Who we are looking for

    We are looking for highly motivated and enthusiastic applicants. Candidates applying for the postdoctoral researcher position are expected to hold a doctoral degree and have a strong academic background, including first-author publications in the fields of plant science, photosynthesis, agriculture, or biotechnology.

    Applicants for the doctoral researcher/project researcher position must hold a Master’s degree (MSc) in biotechnology, plant science, molecular biology, biophysics, or a related field (good academic performance is considered an asset). Students who have not yet completed their degree may also apply and start as research assistants. In such cases, the employment will be temporary until the Master’s degree is completed, after which the title will be updated to project researcher. Once the applicant has received the right to study and has been accepted into the UTUGS doctoral programme, the title will be updated to doctoral researcher.

    Eligibility requirements for these positions are defined in the University of Turku’s regulations: https://www.utu.fi/en/university/organisation. A postdoctoral researcher is required to hold a doctoral degree, and a doctoral researcher/project researcher must hold a Master’s degree.

    If you are applying for the doctoral researcher position, you must apply for the right to pursue doctoral studies (in the spring/autumn 2026 application round) at the University of Turku. Please familiarize yourself with the student information and application criteria of the University of Turku Graduate School (UTUGS) and its doctoral programmes. Note that you must meet the criteria required for the position. Pay particular attention to the English language proficiency requirements. The position requires fluent written and spoken English skills.

    We value collegiality and promote equality, diversity, and inclusivity. We encourage all qualified applicants with different backgrounds to apply for our open positions.

    Benefits

    We offer you responsible and interesting tasks and the opportunity to develop your professional skills in a versatile operating environment. You get a professional and inspiring team to support your work. We take care of your well-being at work and the functionality of your everyday life, e.g. with the help of flexible working hours and our well-being services.

    Please read more about University of Turku as an employee from our Come work with us! pages.

    The European Commission has awarded the University of Turku the right to use the HR Excellence in Research quality label. The label is a token of the University’s commitment to continuous development of the position and working conditions of researchers according to the guidelines set forth in the European Charter for Researchers. 

    The university offers good support and orientation for international hires. Please learn more about the Finnish culture and relocation to Finland:

    The University of Turku is among the top of the world’s universities in most rankings and among the best universities in the world in certain fields. Our high-quality basic research, a multidisciplinary research profile and close cooperation in research infrastructures creates the basis for international top research. Read more about University of Turku in international university rankings.

    The University of Turku has several high-quality research infrastructures required for cutting-edge research. Learn more about the roadmap of research infrastructures and how the research infrastructure and expert teams offer tailored solutions for businesses and organizations.

    Salary and trial period

    The salary is determined by the collective agreement system of the Finnish universities. The salaries are determined in accordance with the university salary system for teaching and research personnel. The salary will be commensurate with qualifications and experience based on the University salary system and will at the beginning of the employment be for a postdoctoral researcher approximately 3600-3800 €/month. If a doctoral student is hired for the position, the starting salary will be approximately 2500-2600 €/month and for a research assistant 2280 eur/month.

    The position has a trial period of six months.

    How to apply

    Applicants should enclose with their application:

    • motivation letter, including statement of motivation, summarizing scientific work and research interest (max 1 page)
    • CV, including education, positions, professional experience, names and contact details of two academic referees (max 2 pages)
    • list of publications (please do not attach your thesis or papers)
    • degree certificate(s) in English or in Finnish

    For further information about the position, please contact: professor Yagut Allahverdiyeva, allahve(at)utu.fi

    With questions about the application process, please contact: HR Specialist Paula Luoma paula.luoma(at)utu.fi  ( during 3.10.2025 – 7.10.2025 please contact hr-tech(at)utu.fi)

    This is an enriching job opportunity in a rapidly evolving field  – we warmly welcome you to join our multicultural working community.

    Doctoral researcher in Optical Spectroscopy, Helsinki

    14.8.2025 | Ajankohtaista

    The University of Helsinki, Department of Chemistry, is seeking a Doctoral Researcher to join the research group of University Lecturer Markus Metsälä, starting November 2025 or as agreed. The position is full-time for a maximum of four years, with a six-month trial period at the start.

    The role is part of the project “Gas Ion Separation Combined with Optical Spectroscopy: Next-Generation Detection of Threat Chemicals”, funded by the Research Council of Finland. The project, in collaboration with VERIFIN (Finnish Institute for Verification of the Chemical Weapons Convention), aims to combine ultra-sensitive photoacoustic spectroscopy with a rapid chemical separation technique, Gas Ion Distillation (GID). The Doctoral Researcher will develop a mid-infrared laser spectrometer using cantilever-enhanced photoacoustic spectroscopy (CEPAS) and integrate it with the GID system for selective and sensitive detection of chemical warfare agents. The University of Helsinki has extensive experience in CEPAS for gas sensing applications.

    Qualifications & Skills

    • Master’s degree in chemistry, physics, or a related field.
    • Prior experience in optical spectroscopy or laser instrumentation.
    • Knowledge of chemical separation techniques or ion mobility spectrometry is advantageous.
    • Ability to obtain or already hold doctoral study rights at the University of Helsinki.
    • Excellent English communication skills, both written and verbal.
    • Motivated and enthusiastic about experimental research and trace gas analysis.

    Responsibilities

    • Design, develop, and optimize mid-IR CEPAS-based laser spectrometer.
    • Collaborate with project partners to integrate the spectrometer with the GID setup.
    • Conduct experimental research and contribute to publications and project reporting.
    • Participate in a multidisciplinary, international research environment and contribute to knowledge exchange.

    Opportunities & Benefits

    • Work and study in a supportive, international, and multidisciplinary environment that values equality, diversity, inclusion, and well-being.
    • Relocation support for international staff.
    • Access to occupational health care, insurance for work-related travel and accidents, sports services, and professional development opportunities.
    • Competitive salary based on Finnish university collective agreements.

    Read more about the application process through the link. Application Deadline: September 29, 2025 (23:59 UTC +3)

    Doctoral Researcher in Biotechnology (DSII) / Väitöskirjatutkija, bioteknologia (DSII) – University of Tampere

    5.8.2025 | Avoimet työpaikat

    https://duunitori.fi/tyopaikat/tyo/doctoral-researcher-in-biotechnology-dsii-vaitoskirjatutkija-bioteknologia-dsii-stsam-19454288?utm_source=loimu&utm_medium=site

    Apply latest 18.8.2025

    Työpaikkakuvaus

    Tampere University and Tampere University of Applied Sciences create a unique environment for multidisciplinary, inspirational and high-impact research and education. Our universities community has its competitive edges in technology, health and society. www.tuni.fi/en 

    The Doctoral School of Industry Innovations (DSII) at Tampere University (TAU) explores research questions that are directly relevant to industry and business. Tampere University hires a Doctoral Researcher in the field of Biotechnology to complete a four-year dissertation project under the joint supervision of a university professor and a company representative. The position is located at the Faculty of Medicine and Health Technology at Tampere University.

    DSII offers Doctoral Researchers a unique opportunity to undertake dissertation research, gain a thorough understanding of product development, tackle real-world business challenges and expand their professional networks.

    The partner company for this position is AMYRA Biotech AG, a Swiss company pioneering innovative digestive enzyme supplements and therapeutics that support the digestive capabilities of the intestinal brush border membrane (BBM). The BBM is one of the body’s primary producers of digestive enzymes. Chronic inflammation, disease, and/or environmental factors can damage the small intestine resulting in brush border enzyme insufficiencies (reviewed by Bonner et. al. 2025) that may compromise digestive function. AMYRA’s Brush Border Enzyme Supplementation Technology (BBEST) is being developed to supplement the digestive capabilities of the small intestine to address general health and nutrition, food sensitivities, and gastrointestinal diseases.

    The purpose of this project is to develop a platform that companies such as AMYRA can leverage to assess their compounds in high-throughput and cost-effective preclinical models.

    This project is part of a larger Enteropathy Research Consortium, led by AMYRA Biotech AG, with the goal of spearheading cross-functional, collaborative research projects investigating how BBM enzyme production varies in health, disorders, and disease. AMYRA is co-supervising 2-3 PhD researchers at universities across Europe, including projects at University of Oxford and the University of Applied Sciences Northwestern Switzerland (FHNW). The PhD student at Tampere University has the opportunity to benefit both academically and professionally from participation in this cross-functional and international collaboration network. 

    Job description

    We are seeking a Doctoral Researcher for a project to develop, optimize, and validate a high-throughput preclinical model of the small intestinal epithelium in conditions of enteropathy. Specifically, induced pluripotent stem cells (iPSCs) will be differentiated into small intestinal epithelial cells (SIECs), and the efficacy of iPSC-SIECs for modelling celiac disease and non-celiac enteropathy will be assessed. iPSC-SIECs will be used to characterize brush border enzyme expression, activity, and nutrient handling, and will be benchmarked against intestinal organoid models and/or patient biopsy sequencing data to validate the use of iPSC-SIECs as clinically relevant disease models. 

    Ultimately, this position will create new knowledge for academia and industry in the fields of digestion, nutrition, and gastroenterology. The goals are:
    To define brush border enzyme insufficiencies and their functional consequences in iPSC-SIEC models of celiac disease.To extend the iPSC-SIEC platform to model non-celiac enteropathies and identify shared patterns of BBM dysfunction.To benchmark iPSC-SIECs against existing intestinal organoid models and/or human tissue to assess their fidelity as enteropathy models. 

    As a Doctoral Researcher you will gain expertise in working with multidisciplinary and international research area in a stimulating environment. 

    The applicant selected for the position will pursue a doctoral degree in Doctoral Programme in Medicine, Biosciences and Biomedical Engineering at the Tampere University. The expected duration of full-time studies is four years.

    The terms of the dissertation work are defined in the Doctoral Researcher’s personal study and supervision plan. The results will be disseminated in scientific peer-reviewed journals, international conferences, doctoral network events and events with industry. Annual progress reports are required. The personal study and supervision plan includes a detailed annual work schedule. 

    In addition to the dissertation work, Doctoral Researchers are expected to teach or perform other duties, such as teaching, related to their studies. These duties may not exceed five (5) per cent of their annual working hours, unless there are justified reasons to the contrary.

    At the university, the work will be supervised by Docent, PhD Kati Juuti-Uusitalo. PhD Sulayman Mourabit and PhD Erin Bonner are the supervisors from Amyra Biotech AG. You will also be supported by practitioners in the steering group, including company representatives. 

    By participating in DSII events, excursions, meetings and training sessions with your fellow doctoral students, you will both get and provide peer support and make your own contribution to the development of DSII.

    Requirements

    You need to have a completed applicable master’s degree in life sciences, biotechnology, cell biology, molecular biology, or biochemistry. High motivation in persuing doctoral degree is very important. 

    Suitable candidate has:

    • Methodological experience in laboratory work, especially aseptic work
    • Has previous experience in cell culture, and preferably in stem cell culture and differentiation
    • Communication skills needed to successfully perform the duties and functions of the position
    • A good command of both spoken and written English

    You also need to have an enquiring mind, enjoy problem-solving and be able to think independently while also being able to work in a team. 

    Please note that before commencing the position, the selected candidate must be accepted to have study rights for the Doctoral Programme in Medicine, Biosciences and Biomedical Engineering (DPMBBE) at Tampere University , unless they already have one. Please visit the admissions webpage for more information on eligibility requirements.

    Tampere University is a unique, multidisciplinary, and boldly forward-looking community. Our values are openness, diversity, responsibility, courage, critical thinking, and learner-centeredness. We hope that you can embrace these values and promote them in your work.

    We offer

    The position will be filled for a fixed-term period of four (4) years, commencing in December 2025 or as soon after that as possible. A trial period of six (6) months applies to all our new employees. Remote work opportunities are agreed separately when needed.

    The salary will be based on both the job requirements and the employee’s personal performance in accordance with the Finnish University Salary System. According to the criteria applied to teaching and research staff, the position of a Doctoral Researcher is placed on level 2-4 of the job requirements scale. A typical starting salary for a Doctoral Researcher is approximately 2700 EUR per month. The salary increases based on experience and the progress of doctoral studies. 

    We invite you to participate a vibrant, active and truly international research community. Our Celiac In Vitro Models research group aims to develop physiologically relevant induced pluripotent stem (iPSC) cell based preclinical models for celiac disease, and other enteropathies. The group is part of Celiac Research Centre.

    Our university has a strong commitment to the highest level of scientific research and the provision of high-quality doctoral education. The faculty as well as the Doctoral School of Industry Innovations support the doctoral students by offering training and peer support. Doctoral students in DSII develop vital skills, enabling them to conduct agile, customer-oriented research, development and business.The University offers a wide range of staff benefits, such as occupational health care, flexible working hours, excellent sports facilities on campus, and several on-campus restaurants and cafés with staff discounts. Please read more about working at Tampere University

    Tampere is one of the major academic hubs in the Nordic countries and offers a dynamic living environment. Tampere region is one of Finland’s three most rapidly growing urban areas and home to a vibrant knowledge-intensive entrepreneurial community. The city is an industrial powerhouse that enjoys a rich cultural scene and a reputation as a centre of Finland’s information society. For more information on Tampere, please visit here.

    How to apply

    Please submit your application through the Tampere University’s online recruitment system. The closing date for applications is 18th of August, 2025 (at 23:59 EEST / UTC+3). Please write your application and all the accompanying documentation in English and attach them in PDF format.

    Please attach only the following documents to your application: 

    • A letter of motivation and description of your research interests (max. 1 A4 page)
    • Curriculum vitae according to the TENK template (including the contact details of two referees).
    • List of publications
    • Degree certificates
    • Transcript of records (master’s degree studies).

    For further information, please contact:

    Senior Reserach Fellow Kati Juuti-Uusitalo, email: kati.juuti-uusitalo@tuni.fi; tel +358 401904292

    Doctoral Researcher in Cell and Molecular Biology – University of Jyväskylä

    5.8.2025 | Avoimet työpaikat

    https://ats.talentadore.com/apply/doctoral-researcher-in-cell-and-molecular-biology/m23L1V

    Apply latest 15.8.2025

    Job description

    The Department of Biological and Environmental Science is seeking to recruit a Doctoral Researcher in Cell and Molecular Biology starting 1st of September 2025 or as soon as possible after that for a maximum of four years. The fixed-term position is for completing doctoral studies.

    Understanding the intranuclear mechanisms of herpesvirus reactivation and its effects on host chromatin structure and dynamics is essential for both basic research and the development of oncolytic virus therapies and novel antivirals that prevent virus reactivation. This PhD project aims to refine our understanding of the mechanisms underlying virus reactivation and the roles of chromatin organization and phase separation in this process. The project is part of an Academy Fellowship funded by the Research Council of Finland, titled ”The role of DNA positioning and interactions in herpesvirus reactivation.” The research will be conducted at the University of Jyväskylä using a multidisciplinary approach that combines cell biology and biophysics with advanced imaging techniques, protein structure prediction algorithms, and biophysical modeling. The project will be carried out in collaboration with the Vihinen-Ranta research group (https://mvr-lab.org) and international partners. Both the research and imaging facilities at the University of Jyväskylä and those of our collaborators in Europe and the USA will be utilized. 

    Who we are looking for

    We are looking for a skilled and motivated PhD candidate with a relevant Master’s degree (e.g., in biochemistry, molecular medicine, or cell and molecular biology). Previous experience in virus research, confocal laser scanning microscopy, and image analysis is considered an advantage. Strong teamwork skills and motivation for scientific research are essential.

    The tasks of a Doctoral Researcher focus on research aiming at the completion of a doctoral thesis, doctoral studies, assisting teaching tasks, and other related tasks to these. The duties, qualification requirements, and the language skills of Doctoral Researchers are stipulated by the University of Jyväskylä Regulations and language skills guidelines (Doctoral Students). A good command of English is required. A trial period of six months will be used in the beginning of the employment.

    The eligible candidate has, or is about to obtain, a suitable Master’s degree. In addition, the Doctoral Researcher must have a permission for post-graduate studies at the University of Jyväskylä. If the applicant does not have permission for post-graduate studies, it must be applied with a separate application procedure from the Faculty of Mathematics and Science after selection for the position.

    Benefits

    At the University of Jyväskylä, you are a recognized member of our community with a unique opportunity to influence international research. You get to participate in our international and multidisciplinary community where the welfare of each individual is important. At the University of Jyväskylä, we offer a great and lively campus area with opportunities to maintain an active and healthy lifestyle.

    Finland has a high standard of living, with free schooling (also in English), affordable childcare, good family benefits, and healthcare. Jyväskylä is located in central Finland in the Finnish lakeland and has excellent opportunities for different nature, outdoor, and sports activities. The city of Jyväskylä is a major educational center and the city has a large student population. As such there is a vibrant cultural scene in the city. To find useful information about the University of Jyväskylä, the City of Jyväskylä and living in Finland, see the University’s International Staff Guide.

    The salary for a Doctoral Researcher is determined based on the task-specific demand level 2-4 of the university salary system for teaching and research staff. The starting salary will be approximately 2400 €/month (gross income). With progress of thesis work, the salary will be revised in accordance with the Collective Agreement of Finnish Education Employers.

    How to apply? 

    The application documents that should be included in English and in PDF format:
    A Curriculum vitae, composed according to good scientific practice and considering the template for researcher’s curriculum vitae by The Finnish Advisory Board on Research Integrity.A short and free-form motivation/cover letter (max 2 pages), including your research interests, career goals and previous experience and merits in relation to the qualifications for this position.Degree certificate (highest degree).

    Applicants are kindly asked to submit their applications using the online application form by 15th of August 2025. Applications sent via email are not considered.

    For further information, please contact Principal investigator, Academy Research Fellow, Salla Mattola, salla.m.mattola@jyu.fi.

    We offer good working conditions for researchers and promote the principles of HR Excellence in Research, HRS4R. The Human Resources Strategy for Researchers (HRS4R) is based on the values of the European Charter for Researchers & The Code of Conduct for the Recruitment of Researchers (Charter & Code). JYU is committed to fairness, consistency and transparency in selection decisions, and undertakes the recommendations for the responsible evaluation of a researcher in Finland. JYU has signed the DORA Declaration. By signing the agreement, JYU is committed to complying with the CoARA principles.

    Our application process is transparent, professional and international. We communicate openly with the applicants throughout the recruitment process. All appointment decisions are based on careful consideration within the faculty. We value equality and diversity in our work community and encourage qualified applicants, regardless of background, to apply for this position.

    Department of Biological and Environmental Science

    The research in the Department of Biological and Environmental Science focuses on three core areas: evolutionary biology, natural resources and environment, and biological nanoscience. The Department building, Ambiotica, is an integral part of the University of Jyväskylä’s Ylistönrinne Mathematics and Science Research Complex, which includes buildings of chemistry, physics and nanoscience. Ambiotica houses high quality laboratories for bioscience and experimental facilities for a wide range of biological and chemical analyses as well as highly motivated laboratory personnel. 

    The University of Jyväskylä is a human-centered environment of 2,500 experts and 14,500 students. Our goal is to create wisdom and wellbeing for all. JYU’s naturally beautiful campus is located in the heart of the city of Jyväskylä. JYU is the third largest employer in Central Finland. 

    MSCA Doctoral Researcher for profiling of metastases of different cancer types – University of Helsinki

    31.10.2024 | Avoimet työpaikat

    https://duunitori.fi/tyopaikat/tyo/msca-doctoral-researcher-for-spatial-multiplexed-immunofluorescence-profiling-of-cancer-types-shsel-18688906

    FIMM is currently seeking a

    MSCA Doctoral Researcher for spatial multiplexed immunofluorescence profiling of metastases of different cancer types

    in a team led by Dr. Teijo Pellinen

    We are seeking a highly motivated Doctoral Researcher for a fully funded, 4-year research position in multiplexed immunofluorescence profiling of metastases from various cancer types. This project, part of the Mac4Me consortium, involves advanced tissue imaging and computational analysis to study immune-tumor cell interactions in clinical samples, including brain, bone, and liver metastases. You will work closely with Genevia Technologies Oy (Finland) and The Norwegian Radium Hospital (Norway) to integrate imaging data with multi-omics datasets using cutting-edge AI tools.

    This position is part of the European Union’s Horizon Europe MSCA Doctoral Network programme, which focuses on identifying new immunotherapy targets through innovative research on tumor-immune system interactions. The Mac4Me network provides a unique training environment, combining expertise from 18 academic and industry partners across Europe. Successful candidates will benefit from interdisciplinary training, research visits, and opportunities for skill development in fields ranging from systems biology to AI-driven cancer research.

    For more information, visit: https://www.van-weerden-lab.com/about

    Work tasks
    As a Doctoral Researcher in Dr. Teijo Pellinen’s team, you will apply innovative multiplexed immunofluorescence technology to spatially profile cells and their interactions in intact tumor tissue sections. You will analyze imaging data to uncover clinically relevant patterns by linking the imaging results to patient health records using advanced computational methods. More information on Dr. Pellinen’s work can be found here (Teijo Pellinen – University of Helsinki).

    Key objectives of this PhD project include:
    1. Designing and constructing tissue microarrays (TMAs) with appropriate control tissues, including tumor tissues (prostate cancer, breast cancer, neuroblastoma), metastasis target tissues (bone, liver, brain), and tissues with high immune cell content (e.g., lymph node, appendix).
    2. Collecting and organizing clinical and pathological data associated with these tissue samples into a shared clinical database.
    3. Developing and optimizing antibody panels for multiplex fluorescence immunohistochemistry (mfIHC).
    4. Performing mfIHC on TMAs and analyzing the resulting high-resolution images for single-cell phenotyping.

    Expected results
    This project will provide crucial insights into immune cell behaviour within metastases from prostate cancer, breast cancer, and neuroblastoma. Through single-cell spatial imaging, you will identify the key immune cell subsets and their activation or inhibition states in clinical metastasis samples. The project will also reveal the interactions and proximities between immune cells and tumor cells, offering a deeper understanding of immune responses in metastatic environments.

    The imaging data will be systematically indexed and integrated with clinical datasets. You will collaborate with partners from Genevia Technologies Oy and The Norwegian Radium Hospital to merge your results with additional multi-omics data. These analyses will contribute to identifying new immune targets and potential therapeutic approaches for cancer treatment.

    The Doctoral Researcher will be based at the Institute for Molecular Medicine Finland (FIMM), University of Helsinki, and work within a vibrant multidisciplinary team embedded in the wider context of Dr. Olli Kallioniemi’s Precision Systems Medicine group at the University of Helsinki (Precision systems medicine | University of Helsinki).

    Opportunities for professional development include attending relevant University of Helsinki courses, presenting your findings at national and international conferences, and conducting research visits to other partners in the Mac4Me Doctoral Network.

    Eligibility criteria
    To be eligible for this position, you must meet the following criteria:
    • Transnational mobility: At the time of recruitment, you must not have resided or carried out your main activity (work, studies, etc.) in Finland for more than 12 months in the 3 years immediately prior to your start date.
    • Early-Stage Researchers (ESRs): You must have less than 4 years of research experience since completing your first MSc degree and must not have been awarded a doctoral degree at the time of recruitment.
    • English proficiency: You must have a strong command of the English language, demonstrated through a recognized English proficiency certificate.

    Please ensure that you meet all these criteria before applying.

    Qualifications

    The ideal candidate will have:

    MSc degree in a relevant field such as molecular biology, biochemistry, biotechnology, bioengineering, computer science, biostatistics, bioinformatics, biomathematics, systems biology, biomedical sciences, or a closely related discipline.
    Strong analytical and communication skills, including proficiency in scientific writing and confident interpersonal communication.
    Experience in bioinformatics or large-scale omics data analysis, including familiarity with commonly used tools and methods.
    Programming skills (e.g., Python, R) are considered an advantage.
    Knowledge of cancer research, preclinical model systems, and image analysis techniques is beneficial but not mandatory.

    We are seeking a motivated and independent researcher who is eager to learn new skills and contribute to a multidisciplinary project at the intersection of cancer biology, immunology, and data science.

    Personal qualities
    We are looking for a candidate who possesses the following personal attributes:
    • Eagerness to learn and develop new skills across multidisciplinary fields.
    • Ability to work both independently and as part of a collaborative team.
    • Problem-solving mindset with a proactive and solution-oriented approach.
    • Strong communication skills, especially when working within a multidisciplinary team.
    • Commitment to meeting deadlines and delivering high-quality results.
    We offer
    • A dynamic and inspiring work environment with access to cutting-edge technology and a vibrant, multidisciplinary research community.
    • Flexible working hours and regular, personalized supervision to support your professional development.
    • Access to unique datasets and cutting-edge methodologies in cancer research, molecular pathology, and image analysis.
    • Competitive salary with yearly wage agreements in accordance with Finnish university guidelines.
    • Comprehensive benefits including pension fund membership and insurance coverage.
    • Opportunities for career development, including attending relevant courses, presenting at national and international conferences, and research visits to partner institutions across Europe.
    • Work-life balance in a supportive and diverse working culture, aligned with the Nordic approach to well-being.

    To apply, please submit the application through the University of Helsinki electronic recruitment system by clicking on Apply for job. 

    Please apply no later than 22nd of November 2024

    The employment may begin as soon as possible or as agreed.

    The University of Helsinki welcomes applicants of any gender and age, linguistic or cultural background, or minority group.
    Click this link to read about accessibility and inclusivity at our University.
    JOIN US TO BUILD A BETTER WORLD – TOGETHER! 
    #HelsinkiUniCareers

    The Institute for Molecular Medicine Finland (FIMM) is an international research institute focusing on human genomics and personalised medicine at the Helsinki Institute of Life Science (HiLIFE) of the University of Helsinki FIMM integrates molecular medicine research, technology centre and biobanking infrastructures under one roof, promoting translational research in grand challenge projects, specifically, the impact of genome information from the Finnish population in personalised health and medicine, individualised cancer medicine, and digital molecular medicine. FIMM is part of the Nordic EMBL Partnership for Molecular Medicine, composed of the European Molecular Biology Laboratory (EMBL) and the centres for molecular medicine in Norway, Sweden and Denmark, and the EU-LIFE Community.